(
  (
    (
      Drosophila_melanogaster,
      Caenorhabditis_elegans
    )
,
    (
      (
        Ciona_intestinalis,
        Ciona_savignyi
      )
,
      (
        (
          Eptatretus_burgeri,
          Petromyzon_marinus
        )
,
        (
          (
            (
              (
                (
                  (
                    (
                      (
                        (
                          (
                            (
                              Astyanax_mexicanus,
                              Pygocentrus_nattereri
                            )
,
                            Electrophorus_electricus
                          )
,
                          Ictalurus_punctatus
                        )
,
                        (
                          (
                            (
                              Cyprinus_carpio_carpio,
                              Sinocyclocheilus_grahami
                            )
,
                            Carassius_auratus
                          )
,
                          Danio_rerio
                        )

                      )
,
                      (
                        Clupea_harengus,
                        Denticeps_clupeoides
                      )

                    )
,
                    (
                      (
                        (
                          (
                            (
                              (
                                (
                                  (
                                    (
                                      (
                                        Kryptolebias_marmoratus,
                                        Nothobranchius_furzeri
                                      )
,
                                      (
                                        (
                                          Fundulus_heteroclitus,
                                          Cyprinodon_variegatus
                                        )
,
                                        (
                                          (
                                            (
                                              Poecilia_latipinna,
                                              Poecilia_formosa
                                            )
,
                                            Poecilia_reticulata
                                          )
,
                                          Xiphophorus_maculatus
                                        )

                                      )

                                    )
,
                                    (
                                      (
                                        Oryzias_javanicus,
                                        Oryzias_melastigma
                                      )
,
                                      (
                                        Oryzias_latipes,
                                        Oryzias_sinensis
                                      )

                                    )

                                  )
,
                                  (
                                    (
                                      (
                                        (
                                          (
                                            (
                                              (
                                                Maylandia_zebra,
                                                Astatotilapia_calliptera
                                              )
,
                                              Pundamilia_nyererei
                                            )
,
                                            Haplochromis_burtoni
                                          )
,
                                          Neolamprologus_brichardi
                                        )
,
                                        Oreochromis_niloticus
                                      )
,
                                      Amphilophus_citrinellus
                                    )
,
                                    (
                                      (
                                        (
                                          Amphiprion_percula,
                                          Amphiprion_ocellaris
                                        )
,
                                        Acanthochromis_polyacanthus
                                      )
,
                                      Stegastes_partitus
                                    )

                                  )

                                )
,
                                (
                                  (
                                    (
                                      Cynoglossus_semilaevis,
                                      Scophthalmus_maximus
                                    )
,
                                    (
                                      (
                                        Seriola_dumerili,
                                        Seriola_lalandi_dorsalis
                                      )
,
                                      Lates_calcarifer
                                    )

                                  )
,
                                  (
                                    (
                                      Anabas_testudineus,
                                      Betta_splendens
                                    )
,
                                    Mastacembelus_armatus
                                  )

                                )

                              )
,
                              (
                                (
                                  (
                                    (
                                      (
                                        Cyclopterus_lumpus,
                                        Gasterosteus_aculeatus
                                      )
,
                                      (
                                        Cottoperca_gobio,
                                        Sander_lucioperca
                                      )

                                    )
,
                                    (
                                      (
                                        Larimichthys_crocea,
                                        Dicentrarchus_labrax
                                      )
,
                                      Sparus_aurata
                                    )

                                  )
,
                                  Labrus_bergylta
                                )
,
                                (
                                  Tetraodon_nigroviridis,
                                  Takifugu_rubripes
                                )

                              )

                            )
,
                            Hippocampus_comes
                          )
,
                          Myripristis_murdjan
                        )
,
                        Gadus_morhua
                      )
,
                      (
                        (
                          (
                            (
                              (
                                Oncorhynchus_kisutch,
                                Oncorhynchus_tshawytscha
                              )
,
                              Oncorhynchus_mykiss
                            )
,
                            (
                              Salmo_salar,
                              Salmo_trutta
                            )

                          )
,
                          Hucho_hucho
                        )
,
                        Esox_lucius
                      )

                    )

                  )
,
                  (
                    Scleropages_formosus,
                    Paramormyrops_kingsleyae
                  )

                )
,
                Lepisosteus_oculatus
              )
,
              Erpetoichthys_calabaricus
            )
,
            (
              (
                (
                  Leptobrachium_leishanense,
                  Xenopus_tropicalis
                )
,
                (
                  (
                    (
                      (
                        (
                          (
                            (
                              (
                                Erinaceus_europaeus,
                                Sorex_araneus
                              )
,
                              (
                                (
                                  (
                                    (
                                      (
                                        Rhinolophus_ferrumequinum,
                                        Myotis_lucifugus
                                      )
,
                                      Pteropus_vampyrus
                                    )
,
                                    (
                                      Equus_caballus,
                                      Equus_asinus_asinus
                                    )

                                  )
,
                                  (
                                    (
                                      (
                                        (
                                          (
                                            (
                                              (
                                                Ovis_aries_reference_breed,
                                                Capra_hircus_reference_breed
                                              )
,
                                              (
                                                (
                                                  (
                                                    Bos_indicus_x_Bos_taurus,
                                                    Bos_taurus
                                                  )
,
                                                  (
                                                    Bos_mutus,
                                                    Bos_grunniens
                                                  )

                                                )
,
                                                Bison_bison_bison
                                              )

                                            )
,
                                            Moschus_moschiferus
                                          )
,
                                          Cervus_hanglu_yarkandensis
                                        )
,
                                        (
                                          (
                                            (
                                              (
                                                (
                                                  Monodon_monoceros,
                                                  Delphinapterus_leucas
                                                )
,
                                                Phocoena_sinus
                                              )
,
                                              Tursiops_truncatus
                                            )
,
                                            Physeter_catodon
                                          )
,
                                          Balaenoptera_musculus
                                        )

                                      )
,
                                      (
                                        (
                                          (
                                            (
                                              (
                                                (
                                                  (
                                                    Sus_scrofa_breed_jinhua,
                                                    Sus_scrofa_breed_meishan
                                                  )
,
                                                  Sus_scrofa_breed_rongchang
                                                )
,
                                                Sus_scrofa_breed_tibetan
                                              )
,
                                              Sus_scrofa_breed_bamei
                                            )
,
                                            Sus_scrofa_breed_wuzhishan
                                          )
,
                                          (
                                            (
                                              Sus_scrofa_reference_breed,
                                              Sus_scrofa_breed_usmarc
                                            )
,
                                            (
                                              (
                                                (
                                                  Sus_scrofa_breed_hampshire,
                                                  Sus_scrofa_breed_berkshire
                                                )
,
                                                Sus_scrofa_breed_landrace
                                              )
,
                                              (
                                                Sus_scrofa_breed_largewhite,
                                                Sus_scrofa_breed_pietrain
                                              )

                                            )

                                          )

                                        )
,
                                        Catagonus_wagneri
                                      )

                                    )
,
                                    (
                                      Camelus_dromedarius,
                                      Vicugna_pacos
                                    )

                                  )

                                )
,
                                (
                                  (
                                    (
                                      (
                                        (
                                          Ursus_maritimus,
                                          Ursus_americanus
                                        )
,
                                        Ailuropoda_melanoleuca_reference_isolate
                                      )
,
                                      (
                                        Neogale_vison,
                                        Mustela_putorius_furo
                                      )

                                    )
,
                                    (
                                      (
                                        Canis_lupus_familiaris_breed_Labrador_retriever,
                                        Canis_lupus_dingo
                                      )
,
                                      Vulpes_vulpes
                                    )

                                  )
,
                                  (
                                    (
                                      (
                                        Panthera_leo,
                                        Panthera_pardus
                                      )
,
                                      Panthera_tigris_altaica
                                    )
,
                                    Felis_catus
                                  )

                                )

                              )

                            )
,
                            (
                              (
                                (
                                  (
                                    (
                                      (
                                        (
                                          (
                                            (
                                              (
                                                (
                                                  Cricetulus_griseus,
                                                  Mesocricetus_auratus
                                                )
,
                                                Peromyscus_maniculatus_bairdii
                                              )
,
                                              Microtus_ochrogaster
                                            )
,
                                            (
                                              (
                                                (
                                                  (
                                                    (
                                                      Mus_spicilegus,
                                                      Mus_spretus_strain_SPRET/EiJ
                                                    )
,
                                                    (
                                                      (
                                                        (
                                                          (
                                                            (
                                                              (
                                                                (
                                                                  Mus_musculus_reference_CL57BL6_strain,
                                                                  Mus_musculus_strain_C57BL/6NJ
                                                                )
,
                                                                Mus_musculus_strain_NZO/HlLtJ
                                                              )
,
                                                              (
                                                                (
                                                                  (
                                                                    (
                                                                      Mus_musculus_strain_A/J,
                                                                      Mus_musculus_strain_BALB/cJ
                                                                    )
,
                                                                    (
                                                                      (
                                                                        Mus_musculus_strain_C3H/HeJ,
                                                                        Mus_musculus_strain_CBA/J
                                                                      )
,
                                                                      Mus_musculus_strain_DBA/2J
                                                                    )

                                                                  )
,
                                                                  Mus_musculus_strain_AKR/J
                                                                )
,
                                                                (
                                                                  Mus_musculus_strain_FVB/NJ,
                                                                  Mus_musculus_strain_NOD/ShiLtJ
                                                                )

                                                              )

                                                            )
,
                                                            (
                                                              Mus_musculus_strain_129S1/SvImJ,
                                                              Mus_musculus_strain_LP/J
                                                            )

                                                          )
,
                                                          Mus_musculus_domesticus_strain_WSB/EiJ
                                                        )
,
                                                        Mus_musculus_musculus_strain_PWK/PhJ
                                                      )
,
                                                      Mus_musculus_castaneus_strain_CAST/EiJ
                                                    )

                                                  )
,
                                                  Mus_caroli_strain_CAROLI_EIJ
                                                )
,
                                                Mus_pahari_strain_PAHARI_EIJ
                                              )
,
                                              Rattus_norvegicus_strain_BN/NHsdMcwi
                                            )

                                          )
,
                                          Nannospalax_galili
                                        )
,
                                        Jaculus_jaculus
                                      )
,
                                      Dipodomys_ordii
                                    )
,
                                    (
                                      (
                                        (
                                          (
                                            Chinchilla_lanigera,
                                            Cavia_porcellus
                                          )
,
                                          Heterocephalus_glaber
                                        )
,
                                        Octodon_degus
                                      )
,
                                      (
                                        (
                                          (
                                            Urocitellus_parryii,
                                            Ictidomys_tridecemlineatus
                                          )
,
                                          Marmota_marmota_marmota
                                        )
,
                                        Sciurus_vulgaris
                                      )

                                    )

                                  )
,
                                  (
                                    Oryctolagus_cuniculus,
                                    Ochotona_princeps
                                  )

                                )
,
                                Tupaia_belangeri
                              )
,
                              (
                                (
                                  (
                                    (
                                      (
                                        (
                                          (
                                            (
                                              (
                                                Pan_troglodytes,
                                                Pan_paniscus
                                              )
,
                                              Homo_sapiens
                                            )
,
                                            Gorilla_gorilla_gorilla
                                          )
,
                                          Pongo_abelii
                                        )
,
                                        Nomascus_leucogenys
                                      )
,
                                      (
                                        (
                                          (
                                            (
                                              (
                                                Cercocebus_atys,
                                                Mandrillus_leucophaeus
                                              )
,
                                              Papio_anubis
                                            )
,
                                            (
                                              (
                                                Macaca_mulatta,
                                                Macaca_fascicularis
                                              )
,
                                              Macaca_nemestrina
                                            )

                                          )
,
                                          Chlorocebus_sabaeus
                                        )
,
                                        (
                                          Rhinopithecus_roxellana,
                                          Rhinopithecus_bieti
                                        )

                                      )

                                    )
,
                                    (
                                      (
                                        (
                                          Cebus_imitator,
                                          Saimiri_boliviensis_boliviensis
                                        )
,
                                        Callithrix_jacchus
                                      )
,
                                      Aotus_nancymaae
                                    )

                                  )
,
                                  Carlito_syrichta
                                )
,
                                (
                                  (
                                    (
                                      Prolemur_simus,
                                      Propithecus_coquereli
                                    )
,
                                    Microcebus_murinus
                                  )
,
                                  Otolemur_garnettii
                                )

                              )

                            )

                          )
,
                          (
                            Dasypus_novemcinctus,
                            Choloepus_hoffmanni
                          )

                        )
,
                        (
                          (
                            Loxodonta_africana,
                            Procavia_capensis
                          )
,
                          Echinops_telfairi
                        )

                      )
,
                      (
                        (
                          (
                            (
                              Vombatus_ursinus,
                              Phascolarctos_cinereus
                            )
,
                            Notamacropus_eugenii
                          )
,
                          Sarcophilus_harrisii
                        )
,
                        Monodelphis_domestica
                      )

                    )
,
                    Ornithorhynchus_anatinus
                  )
,
                  (
                    (
                      (
                        (
                          (
                            (
                              (
                                Notechis_scutatus,
                                Pseudonaja_textilis
                              )
,
                              Laticauda_laticaudata
                            )
,
                            Naja_naja
                          )
,
                          Anolis_carolinensis_reference_strain
                        )
,
                        (
                          Salvator_merianae,
                          Podarcis_muralis
                        )

                      )
,
                      Sphenodon_punctatus
                    )
,
                    (
                      (
                        (
                          (
                            (
                              (
                                (
                                  Gallus_gallus_reference_breed,
                                  Meleagris_gallopavo_reference_strain
                                )
,
                                Coturnix_japonica
                              )
,
                              (
                                Anas_platyrhynchos_platyrhynchos,
                                Anser_brachyrhynchus
                              )

                            )
,
                            (
                              (
                                Aquila_chrysaetos_chrysaetos,
                                Strigops_habroptila
                              )
,
                              (
                                (
                                  (
                                    (
                                      Serinus_canaria,
                                      Taeniopygia_guttata
                                    )
,
                                    Geospiza_fortis
                                  )
,
                                  Parus_major
                                )
,
                                Ficedula_albicollis
                              )

                            )

                          )
,
                          Struthio_camelus_australis
                        )
,
                        Crocodylus_porosus
                      )
,
                      (
                        (
                          (
                            Gopherus_evgoodei,
                            Chelonoidis_abingdonii
                          )
,
                          (
                            Terrapene_carolina_triunguis,
                            Chrysemys_picta_bellii
                          )

                        )
,
                        Pelodiscus_sinensis
                      )

                    )

                  )

                )

              )
,
              Latimeria_chalumnae
            )

          )
,
          Callorhinchus_milii
        )

      )

    )

  )
,
  Saccharomyces_cerevisiae;